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She played Dorothy Gale in The Wizard of Oz in fifth grade, and Veronica Mars.She appeared as Harper in a 13-episode series of short films, It's a Mall World, directed by Milo Ventimiglia, that aired on MTV.That same year, she had her first lead role as Sarah Barton in the drama film Bare, which also premiered at Tribeca.It was written and directed by Natalia Leite and follows a young girl living in a small desert town in Nevada as she becomes romantically involved with a female drifter.Agron said in an interview, "They told me to come back with straight hair and to dress sexier.Later that week, I started work." She and the other cast members were awarded the Screen Actors Guild Award for Outstanding Performance by an Ensemble in a Comedy Series in 2010, and were nominated in the same category the following year.
In 2018, she joined the cast of The Laureate as Laura Riding, alongside Kathy Bates, Tom Hughes, and Laura Haddock.
Agron said in a 2009 interview pertaining to her casting session, "I nearly bailed on my audition for the show.
I was so nervous." Before being cast as Quinn, the show's producers wondered if she appeared too innocent.
The film also starred Patrick Wilson, Lena Headey, and John Cho.
Zipper premiered on January 27, 2015 at the Sundance Film Festival, and was released on August 28, 2015 in a limited release and through video on demand.